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When taken orally, metformin is concentrated in the intestine at levels up to 300 times greater than in the blood, significantly influencing the composition and metabolism of the gut microbiota [16]

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Moreover, given that M cells are the main gateway to transport luminal antigens to the underlying lymphoid tissues and evoke mucosal immune responses, one promising strategy for oral vaccine development is exploring the potential of M cells by M cell-targeting ligands